Evolutionary Developmental Biology by Brian K. Hall PhD, DSc, FRSC (auth.)

By Brian K. Hall PhD, DSc, FRSC (auth.)

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Benrhic, marine, most sedentary and pous, foot attached, some burrow, freeswimming larva; body of adult in 2 shell valves to which adult attached by fleshy pedicle; lophophore as feeding organ (may have internal skeleton); no locomotory organs, no complex sense organs , 330 species The Burgess Shale: different types of animals 20 Group Etymology Bauplan 20. Mollusca L. molluscus, soft 21. Priapulida L. p,riapulus, little pems marine, freshwater and terrestrial, rnodified trochophore larva; body of head, foot and visceral hump covered by mantle which often secretes a calcified shell; complex alimentary canal of a muscular buccal mass, toothed radula, salivary glands, stomach; nervous system of circumoesophageal ring, pedal cords and visceral loops; gills, coelom may be reduced; blood system with propulsive heart, arterial and venous system and haemocoel; haemocyanin as respiratory pigment, 100 000 species marine, benthic, free-living, bilaterally syrnmetrical, unsegmented, wormlike, eversible anterior end, protonephridia with solenocytes, nervous system associated with epidermis, no ganglia, large body cavity, 9 species marine, trochophore larva, unsegmented, bilaterally symmetrical, elongate, worm-like, no chaetae or prostomium, terminal mouth surrounded by tentacles, Ushaped gut, one or rwo metanephridia, 320 species marine, mostly sublittoral, trochophore larva, unsegmented, bilaterally symmetrical; nonretractable proboscis, one pair of chae~ae, pair of anal vesicles, 135 specles marine, freshwater and terrestrial, most free-living, trochophore larva, bilaterally symmetrical, segmented, hydrostatic skeleton, cuticle, chaetae, triploblastic, external circular and internal longitudinal muscles, preoral ganglia and pair of ganglionated ventral nerve cords, nephridia, coe1omoducts, schizocoelic coelom, closed tubular circulatory system, 14 000 species 22.

The thiek legs end in claws. ) Reprodueed from Conway Morris and Whittington (1985) Fossils 0/the Burgess Shale. Anational treasure in Yoho National Park, British Columbia, ' Energy, Mines and Resourees, Canada', reprodueed with the permission of the Minister of Supply and Services Canada, 1990. Shale, but in this case it is soft-bodied arthropods (some 20 different types) rather than trilobites that predominate. Some typical Burgess Shale fossils are Marrella splendens, Canadapsis and Aysheaia.

The paper was authored by two previously unknown Naturalists, Pierre-Stanislas Meyranx and Laurencet, the latter so unknown that his first name(s) is (are) still not known. Impatient with the slow response from the Academie in preseming their study, Meyranx and Laurencet requested that their work be examined by a commission. One of the two commissioners appointed at the regular Academie meeting of 8 February, 1830 was Geoffroy. The basic idea in this paper (Aristotle had posited a similar idea) was of Sepia as avertebrate bent back onto itself with the middle of the abdomen as the fulcrum, the legs attached to the head, and the anus and mouth in alignmem; turning the vertebrate upside down and inside out to 'make it' a cephalopod (Fig.

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